)
)
Fedde's 1901 monograph includes a brief mention of stipules in his Latin description (p. 71):
petiolis ad foliorum insertiones articulatis, saepe articulato nodosis infima parte dilatatis vel subdilatatis et stipulis duobus filiformibus praeditis, rarius carum carentibus.Earlier 19th Century descriptions did not note stipules, although Gray's 1980 description of Berberidaceae indicates (p. 14) "no stipules." And stipules are generally ignored in recent Berberis taxonomies; e.g. Ahrendt 1961, Whittemore FNA. Exceptions supportive of Fedde's work are Wooten's 1898 description of B. haematocarpa, with "stipules laciniate" and Hitchcock & Cronquist 1964, with "rarely stipulate but usually with the base of the petiole dilated."
with rachis ["Blattspindel"] to the leaves inserted articulated, frequently divided with nodes the basal portion of which is broadened ["ein wenig scheidenartig" = sheathlike] or nearly so, and provided with two filiform stipules ["fadenfrmige Nebenblattrudimente"], rarely lacking.
Both our species, and hybrids as well, have stipules which are sheathing and adnate to a clearly dilated petiole base. The stipules are winged with (1-) 3 (-4) teeth, varying in width and length. Generally one or two of the teeth are some 2-3 mm. long and often flexuous. When dry the teeth readily break off.
